LAS VEGAS (AP) — Lincoln Riley had a flashback to three years ago, when he was coaching Oklahoma.

The Sooners were on the verge of tying the game against arch-rival Texas when they made a 33-yard run to the endzone with 3 seconds left on the clock, beating the Longhorns.

Knowing that a field goal was all No. 23 Southern California needed Sunday night, Riley thought back to that play. A run was called for Woody Marks, who ran 13 yards up the middle score with 8 seconds left and defeated No. 13 LSU 27-20.

“I said to someone on the headset, ‘I wouldn’t be surprised if he wrinkles this,'” Riley said. “And of course he did.”

Miller Moss passed for 378 yards and a touchdown, and his 20-yard completion to Kyron Hudson and a subsequent targeting penalty on LSU with 18 seconds left set up Marks’ winning score in the season opener for both teams. It was Marks’ second TD run.

“We’ve worked really hard during the off-season to build the identity of a strong team that really cares about each other,” Moss said.

Moss outpaced LSU’s Garrett Nussmeier, who completed 29 of 38 passes for 304 yards with two touchdowns and an interception on the final drive. Moss was equally efficient, completing 27 of 36 passes in a battle of QBs biding their time after sitting behind the two most recent Heisman Trophy winners, LSU’s Jayden Daniels and USC’s Caleb Williams.

USC, joining the Big Ten Conference for the first time, signaled the Trojans could be a force to be reckoned with with the win in the teams’ first meeting in 40 years after a disappointing 8-5 last year.

“That was some pretty good Big Ten football today,” Riley said with a smile.

And it came against a Southeastern Conference opponent hoping to show it belongs in the College Football Playoff. LSU still has a chance to prove that, but this is the Tigers’ third straight season-opener loss under coach Brian Kelly and their fifth straight overall. LSU also saw its four-game winning streak since last season snapped.

“We didn’t play complementary football,” Kelly said. “But what concerns me the most is the personal fouls, the penalties that are selfish. They’re undisciplined penalties. That comes down to me. We pride ourselves on having a disciplined program, but we clearly didn’t do a good enough job because it affected the game.”

The Tigers accounted for about 60 percent of the crowd, an announced record attendance of 63,969 at Allegiant Stadium, which at times felt like Baton Rouge West. There was some star power, too, with LSU greats Shaquille O’Neal and Daniels, with his Heisman Trophy in tow. Former USC Heisman winners Marcus Allen and Matt Leinart were also in attendance.

Two great catches defined the first half.

Hudson jumped high and back, reaching out with his right hand to somehow bring the ball to his other hand and into his body for a Completion of 24 yards to LSU’s 19-yard line. The play stood after video review, and it led to a 2-yard touchdown run by Marks to put the Trojans up 7-0 four minutes into the second quarter.

LSU’s Kyren Lacy answered with his own acrobatic catch, spinning and getting his right foot just inside for a 19 yard touchdown to tie the game halfway through the period.

Both teams also missed opportunities — the Tigers were denied on downs after a drive to the USC 3, and Trojans kicker Michael Lantz missed a 29-yard field goal to the right as the first-half clock expired. The Trojans could have had an extra 30-40 seconds on that final possession, but Riley elected not to call a timeout before LSU tied the game on a 45-yard field goal by Damian Ramos with 1:08 left.

The game was back-and-forth with the Trojans taking a 20-17 lead with 5:44 left in regulation when Moss found Ja’Kobi Lane for a 28-yard touchdown. A 31-yard field goal by Ramos tied the score but left 1:47 on the clock.

It took too long, as Moss led the Trojans on an eight-play, 75-yard drive to win.

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